It would be interesting to see the website traffic they get as well. The Graun, despite moonies objections, is one of the more popular newspaper websites (along with, ironically, the Mail). The Express however is a pretty poor website, it looks and feels cheap compared to those two, which do at least appear structured and able to engage their readers.
It's not often I will praise the Mail but MailOnline is one of the best websites from a usability and aesthetic point of view. The Express is somewhere near the bottom with the Mirror and the Sun.
Some figures here. I find the mail's traffic levels quite surprising given their circulation figures have remained strong as well according to the wikipedia article, so it's not simply a matter of people choosing a different delivery method. Perhaps their high level of showbiz gossip type articles is paying off in attracting new people who wouldn't be interested in the political stories. Also I do notice the mail quite high in the search listings quite often when I'm looking for information on recent news stories.
The challenge, of course, is turning this traffic into revenue as fewer people buy tradtional newspapers.http://www.guardian.co.uk/media/2011/no ... ic-records